The first endiosymbiosis ended in a great diversification of 
a lineage comprising \textit{Red Algae, Green Algae} and \textit{Land Plants} (terrestrial).
Several Second Enbiosymbioses occurred then: two involving a Red 
Algae and other heterotrophic eucaryotes and giving birth to both Brown 
Algae and Dinoflagellates lineages; another involving a Green Algae and 
a heterotrophic eucaryot and giving birth to Euglens.\\
The interesting with the tree produced (especially from DOGMA) is 
that organisms resulting from the first endosymbiosis are distributed in 
every of the lineage found in the chloroplast genome structure 
evolution: with Red Algae chloroplasts together in one lineage, and 
Green Algae and Land Plants chloroplasts together in antoher lineage; 
while oranisms resulting from secondary endosymbioses are localized in 
the tree: both the chloroplasts of Brown Algae and Dinoflagellates 
representatives are found exclusively in the lineage also comprising the 
Red Algae chloroplasts from which they evolved, while the Euglens 
chloroplasts are related to the Green Algae chloroplasts from which they 
evolved. This make sense in term of biology and history of lineages and 
theories of chloroplasts (and so photosynthetic ability) origins in 
different Eucaryotic lineages.
Interestingly, The sole organisms included that possesses a 
chloroplast (and so a chloroplastic genome) but that have lost the 
photosynthetic ability (being parasitic plants) are found at the base of 
the tree, and not together with its related species phylogenetically, 
meaning that functional chloroplast genes are evolutionnary constrained 
when used in photosynthetic process, but loose rapidly their efficiency 
when not used. They are Cuscuta-grovonii an Angiosperm (flowering plant) 
at the base of the DOGMA Angiosperm-Conifers branch, and 
Epipactis-virginiana also an Angiosperm at the complete base of the tree.
Another interesting result is that land plants that 
represent single sublineage originating from the large and diverse 
lineage of green algae in Eucaryots history are present in two different 
branches of the DOGMA tree, associated with Green Algae, one branch 
comprising the basal grade of land plants (mosses and ferns) and the second 
comprising the most internal lineage of land plants (Conifers and flowering plants). 
But independently of their split in two distinct branches of the DOGMA 
tree, the Land Plants always show a higher number of functional genes in 
their chloroplasts than the green algae from which they emerged, probably meaning that 
terrestrial way of life necessitates more functional genes for an 
optimal photosynthesis than marine way of life. But a more detailed 
analysis of selected genes is necessary to better understad the reasons why?



